KOKO
4 letters
Historical clue formulations
- Gorilla that famously learned to 26-Down — 2025-03-25
- Gorilla who famously learned sign language — 2025-02-16
- Gorilla who was said to have developed the vocabulary level of a three-year-old human — 2024-06-19
- Lord High Executioner in "The Mikado" — 2021-09-14
- Gorilla who was famously taught sign language — 2015-11-18
- Baritone in "The Mikado" — 2015-05-24
- Executioner in "The Mikado" — 2013-12-23
- Gorilla pioneering in sign language — 2013-10-15
- Gorilla skilled in sign language — 2011-12-04
- Gorilla famously taught to use sign language — 2010-10-05
- Famous language-signing gorilla — 2004-03-14
- Clown of renown — 2003-07-31
- "The Mikado" character — 2003-02-19
- The Mikado's Lord High Executioner — 1994-11-21
- "Mikado" executioner — 1994-01-04
